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That's definitely a possibility. I don't believe in burn-in much myself (or at least that it's as pronounced as some people think), but a lot of people experience the effects quite profoundly with Ultrasone. It's possible some of those hearing the Signature Pro as harsh are experiencing the burn-in effect themselves, whatever the cause of the effect may be.
Some of my friends found me turning the volume quite loud when listening to songs.
Using my 15V stepdance, input from the 0404usb, I usually turn past 12pm @low gain
I usually play songs(@the same time do the run-in) at this volume, and sometimes I may find the highs is harsh
At lower volume, say@11am, I am happy and comfortable to listen to the songs, well at that volume I don't think you'll get the full soundstage, but it is very comfortable to listen to music at night time.
Throughout the run-in, (mine has passed 800hours, but I still think the signature pro may need some more time)
I've experienced a few times, harsh highs->not so harsh->soft highs->not so soft->harsh highs...
same thing happened to the bass
Oh, I forgot to mention, I remembered at my 300-500hours time, the signature pro sounds really harsh to me to listen to most of the songs.
At those hours to be frank, I was not so happy with my signature pro
Hope you have another time to try another with some more run-in