The dt990 has a bit more bass punch than the dt770, but the bass is accompanied by very sharp highs (very v-shaped).
The dt770 has almost as much bass as the dt990, but the highs are alot lower, which makes the bass stand out more in the sound.
It is rather difficult to explain without listening to both I think.
And no, I do not have the 80 Ohm DT770, I have the 32Ohm LE. But to be frank, I doubt the difference between the 32Ohm and 80Ohm is massive.
As you say, the ATH-M50x is like a more aggressive dt770. I do not have the ATH-M50x, I only have the originial ATH-M50. But from my understanding the sound of the M50 and M50x is almost identical.
And I agree that the M50 is a more aggressive and v-shaped sound than the DT770. The DT990 is even more aggressive and even more v-shaped than the M50.
Both dt770 and dt990 have recessed mids. If you want something with more mid, try the dt880. It has less bass and less highs, but that gives the mids more space in the sound to shine through.
I love listening to classical music with dt880, and some genres in metal, like funeral doom, sound really good with them.
But normal rock and metal, even though it SHOULD sound better with a neutral headphone with more mid-range, does sound better with a v-shape. The rock just gets boring and non-engaging if the bass and treble isn't boosted just a bit.
To be honest, I doubt you will find what you are looking for in this price range. All v-shaped cans will be lacking in mids, and all cans with great mid will be a bit boring due to the non v-shaped sound.
If you want something that does all of this well, you have to get a headphone with a much larger price-tag. I have heard that the newest beyerdynamic, the DT-1770 might just do this, but I haven't had time to buy it and test it myself.