The Pro 900 has the strongest bass of the three by quite a margin. Its mids are slightly recessed, and its tonal quality is a bit off. Some find its high very harsh, but I found them pleasantly aggressive.
The D2000 take the bass of the Pro 900, tones it down a bit and presents it in a more realistic way. I think its mids and highs are better, but it also depends on the genres you listen to.
While the Pro 900 is a great all-arounder, I find the D2000 to be an even better one. If you only listen to things like Dubstep, though, I'd imagine you'd find the Pro 900 to be much better than the Denon.
I didn't like the DT990 600ohm when I had it, it was aggressive, but very very bright and its bass, while amazing for an open can, isn't in the same ballpark as the bass of the Ultrasone and the Denon.