I am going to attribute the warmth to the iDSD BL which pairs beautifully with the AFC because it warms up the sound a little. When paired with the AFO, it makes it a little warm for my tastes. For me, the AFO seems to pair better with a more neutral amp that is a little cooler than the iDSD. I found that I really, really enjoy the AFO with the HA-2SE. The only downside to the Oppo is that the AFO definitely exposes the ESS glare which I don't mind with many headphones.
In my case, a lot of the warmth could be attributed to the iDSD but only time will tell if it will change with burn in.
I rarely bother with the bass switch on the Oppo as I really dislike it. The bass boost creeps up into the lower mids instead of just the bass; I doubt it would sound good to me with the AFO. When I have tried it in the past with other headphones, I didn't notice a change in the treble.
Maybe what you're hearing is more bass, which in turn makes the treble sound less forward because it doesn't stick out above everything as much.
Maybe a bit too forward isn't the right way to put it. It's not overwhelming or sibilant. I think it's more of the ESS glare that comes through on the AFO. It just makes the treble sound a little "off".
I'm going to agree that the HD650 have never sounded dark to me. I disagree in that I believe they do sound fairly warm to me. I think it's remarkable how similarly they sound to the HD600 when people tend to describe them like they're miles apart.
Right out of the box, the bass extension is very good. Slam and impact are not the strongest just because the frequency response starts to decline below 100HZ.
See this post by MrSpeakers on page 3.