yeah, that's sort of how I felt about the HD650 compared to the HE400i. With the Lyr2 and just the right tubes, I really liked the HD650, but even then it was still relatively even with the HE400i. However, the HE400i was way more versatile amp wise. The HE400i could be at 90% of it's potential straight out of an iPhone, and 99% out of the Schitt Fulla. The HD650 didn't pass 90% until it was setup with the Bifrost 4490 and Lyr2. And even then I'd say the HE400i fought the HD650 to a draw. I think sometimes people on here think a headphone is trash unless it requires a nuclear power generator to sound its best. To me the great genius of the HE400i is it sounds incredible out of almost any well made amp. In a lot of ways I think the HE400i benefits more from being fed a quality DAC than a super powerful amp.
Agree with most of what you are saying, only thing I found is that the HD-650 will scale much more than the HE-400i and pass it but you are way into diminishing returns territory. As you say the beauty of the HE-400i is that it sounds good on almost anything where the HD-650 needs a good supporting cast to achieve what it is capable of.
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I think of the HE560 as a "diagonal" move. This is a term I use for a headphone that is objectively better in many ways, but is also different enough that it also comes down to preferences. I can appreciate that the HE560 has a bit better soundstage layering, better treble micro detail, slightly better bass texture. However, I prefer the sound signature of the HE400i, and it's warmer fuller presentation for most of the music I listen to on a consistent basis.
Same here with the HE-560, yes its does almost everything better than the HE-400i but its like the HD-650, a little more though needs to the rest of your audio chain. And to a point it is also a preference in the sound signature that the individual likes.