The HE400 destroys the X2 in it's presentation of bass. The X2 isn't even within the same stratosphere. Much better texture, depth, control. No contest. the problem with the HE400 is it's wonky transtion from upper mids to treble. That area is a disaster for many, which can ruin the HE400 for many people. The X2 is much better behaved in general, and will translate into a more pleasurable experience overall. You have to look at the total image. The bass on the HE400 is near perfection, IMHO, as is it's lower mids. Everything after...not so much. The X2's bass isn't as good, but is still very, very enjoyable. Paired with it's very good mids, and decent upper range, the X2 offers a more ideal image of sound.
As for the X2 being FOTM like the 990 or whatever. That is wrong on many levels. The 990 has many good things about it, but it's treble is a problem, due to it's super emphasis. The X2 doesn't have that problem, at all. If anything, the x2 is more of a bridge between the 880 and 990 sound, with less treble peakiness than either, and more mids than both. Also a soundstage that has more depth (but less width than the 990).
The 990 is a stellar headphone. FOTM... I think not. All the 990 needs is a simple reduction at 9-10khz, and it offers a stellar sound. The treble is much cleaner than the X2's as well.
What the X2 does so well, is figuring out how to keep everything in very good balance to the ears. It's a joy to listen to, for basically any purpose. Also, depending on your gear's coloration, it can be either warm, or neutral sounding.