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Yeah HE6 had freakin' amazing timbre and such realistic and open 3D imaging capabilities, IMO it sounded shoulder-to-shoulder in terms of naturalness to LCD3s when driven well like out of the Bryston setup at the meet I was at. Downside is it has a fairly pronounced treble, so a lot of us who listened to it kinda agreed that it is not something we'd relax and listen all day long, while the LCD3 (and also the HE400s of course) are.
If HE6s had slightly toned down their treble then I couldn't see any flaw with it given I can drive it well.
It has treble extension out the wazoo, but it doesn't actually have that pronounced treble when really well driven on a speaker amp. Great extension, but linear extension doesn't translate to brightness when in balance with the mids & bass. Though I'm guilty of referring to them as "bright" on occasion when I really mean "with fully extended treble." I tried it on Lyr and didn't find it too treble heavy, but Lyr's one of the more powerful amps already. It doesn't drive it with enough finesse, but I do think it drives it with enough power to keep the FR balanced. The Bryston supposedly drives it, but so far the only three headphone amps that are liked with it reliably are the Mjolnir, HF-6, and Dark Star...and among those only the Dark Star is considered excellent, and even that can be beaten by much cheaper speaker amps supposedly.
Over-pronounced treble is the classic sign of insufficient power on them, so I'm guessing the Bryston may not be a good match overall, even if it otherwise sounded good. Driving them on a modest speaker amp, I don't find it treble pronounced at all, no moreso than HE-400 unless the recording itself is throwing treble left and right (a lot of brass, violins, etc.) And I'm someone very sensitive to treble fatigue. I can't stand K702 for very long without dropping the volume to near-nil. D5k is far more strident and fatiguing for me. But my speaker amp is on the warm side, so that could help. They' have great extension but they're not truly bright. They're rather balanced, and they take EQ/tone controls well enough that you can roll them easily into a laid back can. The beauty of LCD-3 of course is that amazing bass response down to 10Hz, and the fact that you don't don't need to build a containment chamber to house the reactor powering your amp
The trick of HE-6 is to think of them as speakers you wear on your head, not as headphones. I'm glad I got on that train, but I nearly didn't for the same reasons most people don't. Building a speaker rig to drive your headphones is not where most people want their headphone listening to go
That said I listened to HE-400 directly after an HE-6 listening session the other day. I was pleasantly impressed. HE-400 holds its own quite well. Yeah, the timbre's not as perfect, the detail not as nuianced, the bass not as textured, and the soundstage not as open. But it wasn't the stunned kind of "eww, I can't listen to this now" kind of experience. I could have easily listened to the whole album again on HE-400 and not thought of it as horrible. HE-6 may be
more speaker-like, but HE-400 is sufficiently speaker-like on its own.
At $400 + $450 for Lyr (ok, plus tubes, but I won't include that) it's a heck of a kit compared to $1300 + $600-$3.8B for speaker amps and a lifetime supply of uranium fuel to drive it