I am not sure about optimal, but I noticed substantial improvements with around 150-200 hours on them. I probably have about 500 hours on them now and occasionally run them out of my Darkvoice 336se. I have since moved on to many other headphones, but I am always pleasantly surprised by how they sound now.
Personally I haven't found any huge improvements in the 555/595 beyond about 20 hours. It's the 650 that seems to need several hundred hours to reach critical mass.
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