My ZOTL was spec'd for 12AT7 tubes (gain factor 60), but Allan at Berning told me I could use any 12A_7 tube without fear of damage.
I tried a couple 12AX7 tubes (gain factor 100), but they sounded too hot, to the point of distortion.
I tried a couple 5751 tubes (gain factor 70), and they sounded fine, adding treble energy and preserving excellent dynamics.
I tried a couple 12AU7 tubes (gain factor 19), and they sounded beautiful in the mids (especially the Siemens), but bass and dynamics suffered badly, and the music sounded boring.
So you're unlikely to damage your amp (thought I'd aks the manufacturer first), but you may find the lower gain factor has effects other than just making the sound louder or quieter.