Pioneer SE-M521 is the most comfortable headphone that i ever owned. Yes, they use synthetic leather but i am able to wear them for 5-7 hours straight with my glasses on!. They are pretty cheap in Amazon and their performance is stunning (for me)!
In terms of "you will not recognize wearing them", the HD800 is the leader, imo.
But from a satisfying standpoint of comfort, the Empyrian rules them all, of course my opinion.
To be fair, I haven't used a lot of different headphones extensively. However, from the 8ish that I have now and in the past I'd say that the Senn HD 650/6XX with stock pads give me the most comfort. I've heard some complain that the clamping force is too tight, mine are not at all. I have worn them for 5+ hours many times with no discomfort. They are the newer Romanian model if that matters.
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