The higher the input impedance of the current gain stage (in this case the mosfet) the smaller the capacitor (C2 or C4) needs to be, if the input stage has a lower input impedance the tube has to output more current , and in this case the lower frequency waves are limited by the size of the capacitor. I have personally found that the IRF510s can be quite variable in input impedance (before taking into account a mosfets high frequency roll off), so I very much advocate the decision that was made in the PCB version to bulk those coupling capacitors to 220nF, the cost difference is negligible at best and the larger values not going to have a noticeable sound colouring effect over the smaller ones, the difference just isn't great enough. All in all the caps keep the DC on both halves of the amp separated larger ones are more easily going to provide the best frequency response (no need to go over 220nF though).
And yes, these capacitors are non polar, you can install them either way around.
Cheers!